Author contributions
C.D.R., A.A., L.T and R.H.N. designed the study; C.D.R. conducted fieldwork; C.D.R. led all analyses with contributions from S.A.; C.D.R. and S.A. conducted lab work; C.D.R., M.D. and V.X.L. discussed the results. C.D.R. led the writing of the manuscript with contributions from all authors.