hS : The average permuted value of gene diversity within the four geographical regions (Africa, America, Asia, and Europe).
hT : The permuted value of gene diversity across all four geographical regions.
GSt : The permuted value of genetic differentiation among the four geographical regions.
VS: The average observed value of gene diversity within the four geographical regions.
VT: The observed value of gene diversity across all four geographical regions.
NSt: The observed value of genetic differentiation among the four geographical regions.