Authors’ contribution
OKI, ARA, and IKJ conceived and designed the study outline. MTI and MSH setup initial experiments. MTI established the methodology with the assistance of NS, TC and MT. MTI, NS processed the experimental data, interpreted the results and generated the figures. ARA designed the primers for multiplex ARMS-PCR and MSH designed the primers for targeted SNP positions. ARA performed bioinformatics and in silico primer validation. MSH analyzed sequence data. NS, ARA, and MSH wrote the manuscript in consultation with OKI, HMA, IKJ and MTI. HMA and IKJ further reviewed and edited the manuscript extensively. IKJ and MAH supervised the project. Final approval was provided by all authors.