Author contributions
Jun Li and Changxun Xin conceived the project; Changxun Xin designed the experiments; Xiaoyan Wu, Shuo Wang, Jianli Shi, Zhe Peng, Chang Liu, Hong Han performed most of the experiments; Chen Li, Yao Tian, Jiaxin Li, Shaojian Xu contributed materials and participated in discussion; Xiaoyan Wu wrote the manuscript; Jun Li and Fan Zhang supervised the work and edited the final version of the manuscript which was read and approved by all authors.