Funding
The study was partly supported by National Key R&D Program (2016YFD0500708), Major Scientific and Technological Innovation Project (MSTIP)(2019JZZY010720),Shandong Province Modern Agricultural Industry Technology System (SDAIT-08-07), Shandong Province agricultural applications of major innovation projects(SD2019XM003, SD2019XM006), Key R & D Projects in Shandong Province(2019GNC106151), Taishan Scholars Project, Agricultural Science and Technology Innovation Project of Shandong Academy of Agricultural Sciences (CXGC2016B14), Agricultural Science and Technology Innovation Project of Shandong Academy of Agricultural Sciences (CXGC2018E10).