Geographic
variation in the skulls of the horseshoe bats, Rhinolophus
simulator and R. cf. simulator: determining the relative
contributions of adaptation and drift using geometric morphometrics.
Gregory L Mutumi1&2*, David S
Jacobs1*, and Lunga Bam3
1 Animal Evolution and Systematics Group (AES),
Department of Biological Sciences, University of Cape Town, Cape Town
7701, South Africa
2 Life and Environmental Sciences Department,
University of California–Merced, Merced, USA
3 Radiation Science Department, South Africa Nuclear
Energy Corporation, Pretoria, South Africa
Running Head: Selection and drift of bats
Key Words
Diversification, geometric morphometrics, Lande’s model,
micro-evolutionary forces, modularity, neutral evolution, speciation,
* Corresponding Author
E-mail:gmutumi@gmail.com (GLM)
E-mail: