Author Contributions
CRH, DTSH, LPM, SHO, and CLL conceptualized the study, CGH and NWF collected field data for the project and LPM and CLL provided additional data and liaised with external research groups. CRH lead the data analysis with assistance from CGH, DTSH, and JCM. MLC created the hibernacula temperature model. All authors provided critical review and editing to the manuscript.
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