Acknowledgements
We thank Tony Stumbo and Kim Garrett for assistance in the field, Malcolm Reid for advice and assistance with LA-ICP-MS, Ken Miller for assistance preparing maps, Ryan Easton for assisting with document preparation, and various landowners for sampling site access. All sampling was conducted in accordance with local laws and regulations, and with approval from the University of Otago Animal Ethics Committee. This work was funded by the Department of Zoology, University of Otago, Dunedin, New Zealand. During the study, JMA was supported by a University of Otago Doctoral Scholarship.