Acknowledgements
We would like to express our sincere gratitude to Ms. Miwako Sugawara
and Mr. Shinjiro Itoh, and all members in the laboratory of Wildlife
Biology and Medicine, Hokkaido University for their efforts of DNA
analysis. We wish to thank Hatsusaburo Ose and all the members of the
Shiretoko Fishery Productive Association for their generous support. We
thank all the members of the Shiretoko Nature Foundation and South
Shiretoko Brown Bear Information Center for their generous support. We
also thank all the people involved in sample collection. This work was
supported by AIR DO Co., Ltd, Daikin Industries, Ltd., Mitsui & Co.
Environment Fund (R14-0026), Fuji Film Green Fund, a Grant-in-Aid for
Scientific Research from Japan Society for the Promotion of Science
(19K06833), and the Environment Research and Technology Development Fund
(4-1905; JPMEERF20194005) of the Environmental Restoration and
Conservation Agency of Japan. This work was supported by the
World-leading Innovative and Smart Education (WISE) Program (1801) from
the Ministry of Education, Culture, Sports, Science, and Technology,
Japan.