Acknowledgements
We thank Dr. Hanqiang Wang (Shanghai Entomological Museum, Chinese Academy of Sciences) and Huicong Xie (College of Life Sciences, East China Normal University) for helping with providing specimens data. We especially thank Jiazhi Zhang, Sainan Zhang, Tianxu Gao, Wenkai Kou, Ziqi Jin for assistance in specimen access. We thank Emeishan National Nature Reserve, Fenshuiling National Nature Reserve, Guangxi Forestry Bureau, Jinyunshan National Nature Reserve, Jiuwanshan National Nature Reserve, Laojunshan Nature Reserve, Leigongshan National Nature Reserve, Maoershan National Nature Reserve, Nonggang National Nature Reserve, Ruijin Ganjiangyuan Nature Reserve Administration, Shiwandashan National Nature Reserve, Taiyanghe Provincial Nature Reserve, Wenshan Prefecture Forestry and Grassland Bureau and Xishuangbanna Tropical Botanical Garden Chinese Academy of Sciences for their cooperation. This study was sponsored by the Natural Science Foundation of Shanghai (19ZR1416100). The authors declare no conflicts of interest.