8Molecular and Biological Agricultural Sciences, Taiwan International Graduate Program, Academia Sinica and National Chung Hsing University, Taipei, Taiwan
*Corresponding author: Biodiversity Research Cent, Academia Sinica, 128 Academia Road, Sec. 2, Taipei 115, Taiwan. Tel : +886-2-2787-1585, Fax: +886-2-2785-8059, e-mail: ryujimachida@gmail.com
#FMGM, CLF, ND, and YHW should be considered the joint first authors.