Hindcast species distribution models projected to the past (Supplementary Data Figure S6) support a long-term presence of D. orientalis and DC2 clades in the eastern Mediterranean region, and a prevalence of the DC3 clade in the western Mediterranean since the Last Interglacial (LIG, ~120,000 - 140,000 years BP). The overlap between current and Mid Holocene (MH, about 6000 years ago) distribution models was high for DC2 (0.907), D. orientalis(0.798) and DC3 (0.859), slightly lower between MH and the Last Glacial Maximum (LGM, about 22,000 years ago) for DC2 (0.835), D. orientalis (0.564) and DC3 (0.719), and moderately to drastically lower between LGM and LIG for DC2 (0.216), D. orientalis (0.419) and DC3 (0.558).