Acknowledgements
This research utilised Queen Mary’s Apocrita HPC facility, supported by QMUL Research-IT http://doi.org/10.5281/zenodo.438045. We thank Phillip Howard, Dr. Chloe Economou for their technical wet lab support, Mason Connolly for his sterling contribution to field collection and preliminary data generation, James Crowe for Bioinformatics support, and Dr. Alex Twyford for insightful comments on the manuscript. Special thanks to Michel and Diana Rey for their hospitality over multiple years to multiple field expeditions. Hannes Becher was supported by a QMUL college studentship.