Author Contributions

The initial camouflage and 3D structure approach was conceived by JT and AH and carried out by GRAH. Colour-calibrated photographs and 3D scans were collected by GRAH, with nests located and monitored by LG and RB. The code for converting .ply files to depth maps and for measuring the difference of Gaussian energy was created by JT and modified by GRAH. GRAH created the code for measuring occlusion. Manuscript and supplementary guides were first written by GRAH, with subsequent edits by all authors. GRAH performed data analyses with guidance from JT and IC.
Acknowledgements.
NERC GW4+ NE/S007504/1 funded GRAH in a CASE partnership with the GWCT. JT was funded by a NERC Independent Research Fellowship NE/P018084/1. We thank the GWCT, and the landowners, gamekeepers and farmers involved associated farming bodies for allowing us to carry out our research at their field sites.
Conflict of Interest
The authors declare no conflict of interest.
Data Accessibility
[Data will be accessible from Dryad on publication, see attached supplementary R folder, temporary link via dropbox https://www.dropbox.com/s/e9kuk9g3x60si2r/R_Code_EE.zip?dl=0], ImageJ scripts for running RNL and 3D analyses with the MICA toolbox can be downloaded from our GitHub, https://github.com/GeorgeHancock471/3D_RNL_Tools.