ACKNOWLEDGMENTS
The project was supported by National Natural Science Foundation of China (32100193,31761133020, 31821001), the Strategic Priority Research Program of Chinese Academy of Sciences (XDB31000000), the Start-up funds for Introduced Talent at Lanzhou University (561119209), Fundamental Research Funds for the Central Universities (lzujbky-2020-ct02) and National Key R&D Program of China (2016YFC0503200). The authors thank Drs. Huang Chengming, Zhou Jiang, Meng Tao, Zhou Qihai and Liu Zhang Liye for their assistance in the collection of field data and comments. We acknowledge Chrissie, Sara, Jenni, and Dax for their support and guidance.