Author Contribution
M.L., A.G. and A.R. planned the labeling experiment; M.L. and A.W. designed the treatment and sampling experiment; A.W. and M.M.L conducted the field work; A.W. and Z.D. conducted experiment work and analyzed the data; A.W. wrote the manuscript; M.L., A.G., A.R., M.M.L and M.S. revised the manuscript